Molecular Genetics and Pathogenesis of Melanoma

Melanoma, a malignancy of melanocytes, is a genetically diverse tumor influenced by UV exposure and other mutational drivers. This blog explores the molecular genetics, subtypes, and clinical implications of melanoma, alongside histological and pathway-focused tables.


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Types of Melanoma Based on UV Damage

1. High-CSD Melanoma (Cumulative Sun Damage)

High-CSD melanomas arise in chronically sun-exposed areas and exhibit features of accumulated UV damage. Key subtypes include:

  • Lentigo Maligna Melanoma
  • High-CSD Nodular Melanoma
  • Desmoplastic Melanoma

2. Low-CSD Melanoma (Minimal UV Damage)

Low-CSD melanomas, such as Superficial Spreading Melanoma, are linked to intermittent sun exposure, often on less chronically exposed areas.


3. No-CSD Melanoma (Sun-Shielded Sites)

These melanomas occur on areas shielded from UV radiation. Key examples include:

  • Mucosal Melanoma
  • Acral Melanoma
  • Melanomas arising in congenital or blue nevi
  • Uveal Melanoma

Types of Melanoma Based on UV Damage

Table 1: Melanoma Types Based on UV Damage

Type of MelanomaUV ExposureSubtypesKey Features
High-CSD MelanomaChronic Sun DamageLentigo Maligna Melanoma, High-CSD Nodular Melanoma, Desmoplastic MelanomaOccurs in areas with chronic UV exposure, high tumor mutational burden (TMB).
Low-CSD MelanomaIntermittent Sun DamageSuperficial Spreading MelanomaLinked to intermittent sun exposure, moderate TMB.
No-CSD MelanomaNone (Sun-Shielded Sites)Mucosal Melanoma, Acral Melanoma, Malignant Spitz Tumor, Uveal MelanomaLow TMB, associated with congenital or site-specific genetic mutations.

Molecular Genetics of Melanoma

MAPK Pathway and Driver Mutations

Melanomas frequently harbor mutations in the MAPK signaling pathway, which drives cellular proliferation:

  • BRAF Mutations (~50%): Predominantly V600E mutation, targeted by inhibitors like vemurafenib.
  • NRAS Mutations (~25%): Linked to aggressive tumor behavior.
  • KIT Mutations (~2%): Found in acral and mucosal melanomas.

Uveal Melanoma

This unique melanoma subtype is driven by mutations in:

  • GNAQ and GNA11: These mutations activate G-protein signaling, leading to proliferation.
  • BAP1: Loss of this tumor suppressor gene is associated with metastasis, particularly to the liver.

Sun Exposure and Tumor Mutational Burden (TMB)

The degree of UV damage correlates with the tumor mutational burden:

  • High-CSD Melanomas: Exhibit high TMB, reflecting chronic UV-induced DNA damage.
  • No-CSD Melanomas: Have very low TMB and are driven by other genetic events, such as congenital mutations.

Molecular Genetics of Melanoma

Table 2: Genetic Mutations in Melanoma by Site

SiteKey Genetic MutationsAssociated Features
CutaneousBRAF (~50%), NRAS (~25%), KIT (~2%)MAPK pathway activation, targeted therapies like BRAF inhibitors.
MucosalKIT mutations, NRAS mutationsRare, occurs in sun-shielded areas, aggressive behavior.
AcralKIT mutations, NRAS mutationsFound on palms, soles, and under nails, not UV-related.
UvealGNAQ, GNA11 (>90%), BAP1 lossDistinct pathway, high metastatic potential, liver metastasis common.
Congenital NeviNRAS mutations, occasional BRAF mutationsPresent at birth, may transform into melanoma.
Blue NeviGNAQ, GNA11Low-risk, can progress to melanoma in rare cases.

Clinical Relevance and Therapeutic Implications

Understanding melanoma’s molecular genetics has revolutionized its diagnosis and treatment:

  1. Targeted Therapy: BRAF and MEK inhibitors provide precision treatment for patients with BRAF mutations.
  2. Immunotherapy: High mutational burden in UV-damaged melanomas makes them suitable candidates for immune checkpoint inhibitors.
  3. Prognostic Insights: NRAS-mutated melanomas have a poorer prognosis compared to BRAF-mutated melanomas.

Table 3: Precursor Lesions of Melanoma and Their Respective Mutations

Precursor LesionAssociated Melanoma TypeCommon Genetic MutationsKey Features
Congenital NevusMelanoma arising in congenital nevusNRAS (~80%), occasional BRAFPresent at birth; larger lesions have a higher risk of melanoma transformation.
Dysplastic NevusSuperficial Spreading MelanomaBRAF (~50%)NRAS (~25%)Atypical nevus with architectural disorder; often associated with familial melanoma syndromes.
Blue NevusMelanoma arising in blue nevusGNAQGNA11Heavily pigmented, spindle-shaped melanocytes; rarely transforms into melanoma.
Atypical Spitz Tumor (Spitz Nevus)Malignant Spitz TumorHRAS, rare BRAF or TERT promoterSpitzoid melanocytes with atypia; rare progression to melanoma.
Nevus SpilusMelanoma arising in nevus spilusNRAS, rare BRAFSpeckled pigmented lesion; may develop melanoma within the nevus.

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References

  1. Robbins and Cotran Pathologic Basis of Disease (10th Edition) – Vinay Kumar, Abul Abbas, Jon Aster.
  2. Weedon’s Skin Pathology (5th Edition) – David Weedon.
  3. Diagnostic Histopathology of Tumors (5th Edition) – Christopher Fletcher.
  4. Molecular Pathology of Melanoma (Springer Series) – Klaus Busam.
  5. Pathology of Melanocytic Tumors (2nd Edition) – Raymond Barnhill.

Disclaimer: This blog is a simplified educational guide.


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